With more than 4,000 entries from 68 countries in this year's Comedy Wildlife Photography Awards, competition was fierce but from 40 finalists, the winners have now been announced.

Overall winner, Sarah Skinner, won for her image entitled 'Grab life by the...' taken of a playful cub in Botswana. Commenting on her winning, Sarah says: “I am absolutely delighted to be awarded the title as Overall Winner in the Comedy Wildlife Photography Awards 2019. It certainly warms my heart to know that this image will spread some laughter and happiness around the world.

"I am happy to report that this lioness continues to thrive in the pride, having seen her again in October this year. I can only hope and encourage everyone, as a collective, to each do our part in the conservation of all wildlife species, so thatfuture generations can enjoy them in the same way that I have done during my career as a wildlife photographer. Long may lions walk the plains...” As her prize, Sarah will be enjoying a safari in the Masai Mara, Kenya, trophy from the Wonder Workshop in Tanzania, a ThinkTank photography bag and iPad preloaded with Affinity Photo software.

Sarah's image also scooped her the Creatures of the Land award.

Here are some of the other highly commended and winning entries, including The Affinity Photo People's Choice Award, voted for by the public. The image by Harry Walker of a surprised otter also scooped the Olympus Creatures Under Water Award.